Undergraduate

Creighton University, Omaha, Nebraska, B.S. (Biology), December 1979

Graduate

Florida State University, Tallahassee, Florida

• M.S., Biological Science (Genetics & Cell Biology), December 1982
• Ph.D., Biological Science (Genetics & Cell Biology), December 1987

Postdoctoral

University of Michigan Medical School, Ann Arbor, Michigan

• Research Associate, Howard Hughes Medical Institute, November 1987 – February 1989
• Postdoctoral Fellow, Dept. of Internal Medicine, March 1989 – August 1991