Undergraduate
Creighton University, Omaha, Nebraska, B.S. (Biology), December 1979
Graduate
Florida State University, Tallahassee, Florida
- M.S., Biological Science (Genetics & Cell Biology), December 1982
- Ph.D., Biological Science (Genetics & Cell Biology), December 1987
Postdoctoral
University of Michigan Medical School, Ann Arbor, Michigan
- Research Associate, Howard Hughes Medical Institute, November 1987 – February 1989
- Postdoctoral Fellow, Dept. of Internal Medicine, March 1989 – August 1991